Most of the time, higher tier monsters have all the abilities of preceding tiers, with increased intensity.
Wolver Traits by Tier
Tier 1
Alpha Wolvers can dash while their underlings lack special abilities.
Tier 2
The Wolvers are more agile and can now dash near players and attack quickly. Alpha Wolvers can burrow and therefore effectively teleport around the room.
Tier 3
All Wolvers can burrow (teleport).
Strategy
Guide: Wolver Combat
The Alpha Wolvers' three-hit attack is quite dangerous and harder to disrupt than most, often requiring multiple hits to interrupt, especially if your weapon is below their level (as will always be true in Danger Rooms due to scaling rules). It is often safest to back up and counter-attack after their combo is finished. Additionally, while they are focused on you, your allies can attack them from behind with impunity.
Many Cutter/Striker users tend to never use their charge attack on this family. However, if for any reason you want to, It is prefered that you charge attack just out of reach of the sword, letting the wolver come into the swords range.
Wolvers occasionally sit and bark. During this time, they do not dodge and any hit will knock them down. Alpha Wolvers bark as well, but won't be knocked down during this time.
If you can mange to pin a Wolver or Alpha Wolver in a corner, attacking continuously with a sword will usually prevent them from attacking or escaping, and you can finish them off easily.
Tip: If you are in combat with a group of wolvers I advise running in circles around them shooting them with your handgun. This will make the have a hard time hitting you, but allow you to hit them.
Wolver are considered Grunt class, Alpha Wolvers and Perma-Frostifur are considered Brute class.
Wolvers are usually found in groups and can call out to their comrades to ambush an unwary adventurer.
As knights progress through the tiers, Wolvers become more agile and move more quickly, turning capable of dodging projectiles directed at them.
Eventually the Wolvers will burrow underground and pop out, preparing to attack a knight from behind.
Wolvers aside from their inherent status are less resistant to Stun and more resistant to Freeze.
According to GM Coriolis, Wolvers appear to be particularly fond of biscuits but apparently still tend to bite the knight that feeds them! The fact was found in relation to a bug in which Tier 3 Wolvers can dig underground and exit the boundaries of the map, the phrase "Bad wolver, no biscuit!" was stated in response to the glitch.
History
This monster has been encountered since release 2011-04-04 when the game officially launched.
List
Wolver
Monstrous Profile
Tier 1
Tier 2
Tier 3
Description
Small wolf-like monsters that are rarely found alone. They would try to sneak upon their prey to take a bite out of them.
Notes
This monster behaves like the others. See above strategy for combat and equip effective gear.
Combat
Elite Difficulty:
Bite
Advanced and Normal Difficulty:
Bite
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
The leaders of various Wolver packs, moving around areas quicker than the rest of their pack.
Notes
This monster behaves like the others. See above strategy for combat and equip effective gear.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Ash covered Wolvers that leave soot everywhere they trot, making them hard to clean up after.
Notes
Like other Fire themed monsters, this enemy is immune to fire. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
Elite Difficulty:
Bite
Advanced and Normal Difficulty:
Bite
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Fast, swift ash covered Alphas that cause can cause a severe burn if bitten.
Notes
Like other Fire themed monsters, this enemy is immune to fire. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Ice coated Wolvers with chilling, razor-sharp teeth.
Notes
Like other Freeze themed monsters, this enemy is immune to frezee. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
Elite Difficulty:
Bite
Advanced and Normal Difficulty:
Bite
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Cold-hearted hunters of the tundra regions within the clockworks.
Notes
Like other Freeze themed monsters, this enemy is immune to frezee. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Sickly Wolvers with a bad case of hydrophobia. Although ailing, they're just as vicious as their cohorts, but they'll occasionally pause to froth a bit.
Notes
Like other Poison themed monsters, this enemy is immune to poison. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
Elite Difficulty:
Bite
Advanced and Normal Difficulty:
Bite
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Deadly disease infested Alphas that have more than just fleas keeping them company.
Notes
Like other Poison themed monsters, this enemy is immune to shock. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Statically charged Wolvers that have a tendency to shock whoever they take a bite of.
Notes
Like other Shock themed monsters, this enemy is immune to poison. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
Elite Difficulty:
Bite
Advanced and Normal Difficulty:
Bite
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Super charged Alphas that have a shocking jaw force and a huge appetite!
Notes
Like other Shock themed monsters, this enemy is immune to shock. It behaves like the others. See above strategy for combat and equip effective gear.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Encounter
Random Spawn
This monster has a very high chance to spawn in these areas:
Alpha Frostifurs that seem to be under the influence of the Permafrost's power and are invulnerable to damage.
Notes
Perma-Frostifur was introduced with release 2012-08-01 along with Dangerous Mission: Heart of Ice. There are only two of these type of Wolvers that accompany Maulos in Amu-Sol.
Combat
3 Successive Bites
Attack Buff (affects all Wolvers)
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Loot
This monster does not drop loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
This dark monster is Swarm-Infested, so it offers absolutely no Crowns, Heat, Materials, or Vials, and seemingly never stops spawning.
Notes
It's a wolver that entered the dark side! Their defenses are enhanced on a Swarm Seed.
Combat
Bite
Dash
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
This monster has not been seen by players. It is however, viewable in the Promotional video of Spiral Knights.
Notes
This monster was in the Preview Event, it has not been seen since.
Combat
Unknown
Loot is dropped when the monster is defeated. The loot that drops is random. Sometimes loot isn't dropped at all. This list is simply a pool of possible loot.
Loot
This monster does not drop loot.
Random spawns also apply to missions with floors that have geography similar to the relevant arcade area. The normal version of a monster can sometimes spawn instead of the status version in areas themed with a status, unless the spawn is guaranteed. See Gate Maps for details.
Stun and more resistant to 
Freeze.
